Joseph Treloar Wearn

University of Pennsylvania; proposed and carried out glomerular micropuncture with A. N. Richards

Dates: 1893-1984 (source: Life dates as given in research/02_uroscopy_and_urine.md §5.7 (graded VERIFIED with the NAS memoir, Wikipedia and historyofmedicine.com) )
Chapters: V. What the Water Knew

What the film says

With Richards, drew fluid from the capsule of a single glomerulus in a frog's kidney.

Watch 24:53 in the film V. What the Water Knew

December 1920: In Philadelphia the team watches Robert Chambers demonstrate micromanipulation; Wearn proposes using it to draw fluid from the glomerular space. timeline
December 1921: Wearn and Richards give a preliminary report to the American Physiological Society; the frog-to-mammal objection is raised and Richards's reply wins an ovation. timeline
1924: Wearn and Richards publish their observations on the composition of glomerular urine in the American Journal of Physiology. timeline
1925: Wearn and Richards report chloride concentrations in frog glomerular urine. timeline

Schmidt 1971, NAS Biographical Memoir of Alfred Newton Richards Secondary
Wearn JT, Richards AN. Observations on the composition of glomerular urine, with particular reference to the problem of reabsorption in the renal tubules. Am J Physiol 1924;71(1):209-227. doi:10.1152/ajplegacy.1924.71.1.209 Primary